Étienne-Jules Marey was a pioneer of fast photographic techniques that he used to investigate movement. In his most iconic photographic studies he took multiple successive images by overlapping celluloid prints on top of one another, which allowed him to see flight phases of birds and study their relations to each other in a single picture. The following video references his famous chronophotographs of birds flying, though in this case the “flight” is carried out by the light pulse.
Motion Study II: Albatross
Museum of Vertebrate Zoology, UC Berkeley